Stephen G. Tilley   
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Stephen G. Tilley is a member of the Department of Biological Sciences at Smith College, Northampton, MA. He has been doing research on and photographing plethodondid salamanders for over 40 years. Address: Department of Biological Sciences, 44 College Lane, Smith College, Northampton, MA 10163. Email:
Special interests: Systematics of dusky salamanders (Genus Desmognathus) in the southern Appalachian Mountains.
Total Photos Contributed: 71  [View all photos] or [See a list of all photos] or [View most recent photos]
Collection:  Private
Email address for Stephen G. Tilley:  stilley[AT]   (Replace the [AT] with the @ symbol before sending an email.)
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